phd programs for biology majors

Best Biological Sciences Programs

Ranked in 2022, part of Best Science Schools

A graduate degree in biological sciences gives students

A graduate degree in biological sciences gives students a broad base of knowledge in the life sciences. The discipline also offers many specialties, such as cell or molecular biology. These are the best schools for biological sciences. Each school's score reflects its average rating on a scale from 1 (marginal) to 5 (outstanding), based on a survey of academics at peer institutions. Read the methodology »

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Biology Graduate Programs in America

1-25 of 398 results

MIT School of Science

• Cambridge, MA ·
• Massachusetts Institute of Technology ·
• Graduate School

Massachusetts Institute of Technology, Graduate School, CAMBRIDGE, MA.

Stanford University Department of Humanities and Sciences

• Stanford, CA ·
• Stanford University ·

Stanford University, Graduate School, STANFORD, CA.

Stanford University School of Medicine

• · Rating 5 out of 5   3 reviews

Stanford University, Graduate School, STANFORD, CA. 3 Niche users give it an average review of 5 stars. Featured Review: Doctoral Student says I begin in the Fall of 2023--Based on my interview, and pre-classes I have taken this program is amazing. It is very student-centered. Read 3 reviews.

Tarleton State University

• Graduate School ·
• STEPHENVILLE, TX
• · Rating 4.68 out of 5   62

Florida Institute of Technology

• MELBOURNE, FL
• · Rating 4.59 out of 5   73

Mississippi College

• CLINTON, MS
• · Rating 4.37 out of 5   38

Harvard Graduate School of Arts and Sciences

• Harvard University ·
• · Rating 4.56 out of 5   9 reviews

Harvard University, Graduate School, CAMBRIDGE, MA. 9 Niche users give it an average review of 4.6 stars. Featured Review: Other says I am Harvard Extension School student pursuing a master degree, ALM, in sustainability. I have achieved a 3.89 in this program so far and have qualified, applied, and accepted as a 'Special Student'... Read 9 reviews.

Harvard Medical School

• Boston, MA ·
• · Rating 4.95 out of 5   19 reviews

Harvard University, Graduate School, BOSTON, MA. 19 Niche users give it an average review of 4.9 stars. Featured Review: Doctoral Student says I have really felt that the curriculum at Harvard is designed with a lot of care by faculty. Each of our block faculty leads are clearly incredibly passionate, and tailor the curriculum to maximize... Read 19 reviews.

Duke University School of Medicine

• Durham, NC ·
• Duke University ·
• · Rating 4.76 out of 5   17 reviews

Duke University, Graduate School, DURHAM, NC. 17 Niche users give it an average review of 4.8 stars. Featured Review: Doctoral Student says Professors are incredible, always willing to go the extra mile to help students succeed. Curriculum is designed to prepare students to be successful therapists. This program is truly incredible. Read 17 reviews.

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Brown University Graduate School

• Providence, RI ·
• Brown University ·
• · Rating 5 out of 5   4 reviews

Brown University, Graduate School, PROVIDENCE, RI. 4 Niche users give it an average review of 5 stars. Featured Review: Master's Student says This is my first semester in the biotechnology program. The program manager is helpful and commutative. I have found a great lab match for my research interests. My classes are interesting. I have... Read 4 reviews.

School of Arts & Sciences - University of Pennsylvania

• Philadelphia, PA ·
• University of Pennsylvania ·

University of Pennsylvania, Graduate School, PHILADELPHIA, PA.

Weinberg College of Arts and Sciences

• Evanston, IL ·
• Northwestern University ·

Northwestern University, Graduate School, EVANSTON, IL.

Vanderbilt University School of Medicine

• Nashville, TN ·
• Vanderbilt University ·
• · Rating 5 out of 5   2 reviews

Vanderbilt University, Graduate School, NASHVILLE, TN. 2 Niche users give it an average review of 5 stars. Featured Review: Graduate Student says Vanderbilt University School of Nursing. Loved my experience! Great teachers, easy to access, clear cut goals in class, great clinical experiences. Read 2 reviews.

Washington University in St. Louis - Arts & Sciences

• St. Louis, MO ·
• Washington University in St. Louis ·

Washington University in St. Louis, Graduate School, ST. LOUIS, MO.

Guarini School of Graduate and Advanced Studies

• Hanover, NH ·
• Dartmouth College ·
• · Rating 5 out of 5   1 review

Dartmouth College, Graduate School, HANOVER, NH. 1 Niche users give it an average review of 5 stars. Featured Review: Alum says I had a beautiful life-changing experience at the grand Guarini School of Graduate and Advanced Studies. The Guarini graduate program MALS created so many beautiful opportunities in interdisciplinary... Read 1 reviews.

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Division of Biology and Biological Engineering - California Institute of Technology

• Pasadena, CA ·
• California Institute of Technology ·

California Institute of Technology, Graduate School, PASADENA, CA.

College of Science - University of Notre Dame

• Notre Dame, IN ·
• University of Notre Dame ·

University of Notre Dame, Graduate School, NOTRE DAME, IN.

Biological Sciences Division - University of Chicago

• Chicago, IL ·
• University of Chicago ·
• · Rating 4 out of 5   1 review

University of Chicago, Graduate School, CHICAGO, IL. 1 Niche users give it an average review of 4 stars. Featured Review: Master's Student says So far so good. Nothing out of the ordinary. Professors are knowledgeable. I would say to ensure that you know your interests and work closely with the advisors to pick suitable classes. Read 1 reviews.

Pritzker School of Medicine

University of Chicago, Graduate School, CHICAGO, IL.

Dornsife College of Letters, Arts and Sciences

• Los Angeles, CA ·
• University of Southern California ·

University of Southern California, Graduate School, LOS ANGELES, CA.

Cornell University College of Arts & Sciences

• Ithaca, NY ·
• Cornell University ·

Cornell University, Graduate School, ITHACA, NY.

Georgetown University School of Medicine

• Nw Washington, DC ·
• Georgetown University ·
• · Rating 4.75 out of 5   4 reviews

Georgetown University, Graduate School, NW WASHINGTON, DC. 4 Niche users give it an average review of 4.8 stars. Featured Review: Doctoral Student says Compressed pre-clinical work. Administration has been working on finding better and better educators, as well as hiring more learning specialists for the student body. Could have more focus on Step 1... Read 4 reviews.

School of Arts and Sciences - Tufts University

• Medford, MA ·
• Tufts University ·
• · Rating 4 out of 5   4 reviews

Tufts University, Graduate School, MEDFORD, MA. 4 Niche users give it an average review of 4 stars. Featured Review: Master's Student says My experience was mostly great! I felt supported by faculty and staff and became involved in student activities. It was just unfortunate that my on-campus experience was cut short due to the pandemic... Read 4 reviews.

UCLA College of Letters and Science

• University of California - Los Angeles ·
• · Rating 3 out of 5   1 review

University of California - Los Angeles, Graduate School, LOS ANGELES, CA. 1 Niche users give it an average review of 3 stars. Read 1 reviews.

Krieger School of Arts & Sciences

• Baltimore, MD ·
• Johns Hopkins University ·
• · Rating 4.52 out of 5   21 reviews

Johns Hopkins University, Graduate School, BALTIMORE, MD. 21 Niche users give it an average review of 4.5 stars. Featured Review: Master's Student says Starting this fall, my graduate journey is set to begin on an exciting note. I’ve already connected with two professors who have proven to be incredibly supportive and kind. I can't wait to dive into... Read 21 reviews.

Mellon College of Science

• Pittsburgh, PA ·
• Carnegie Mellon University ·

Blue checkmark. Carnegie Mellon University, Graduate School, PITTSBURGH, PA.

The Graduate School of Arts & Sciences - University of Virginia

• Charlottesville, VA ·
• University of Virginia ·

University of Virginia, Graduate School, CHARLOTTESVILLE, VA. 1 Niche users give it an average review of 4 stars. Featured Review: Alum says Very good in some areas, excellent in other areas, many academic choices available in all areas of study Read 1 reviews.

College of Sciences - Georgia Institute of Technology

• Atlanta, GA ·
• Georgia Institute of Technology ·

Georgia Institute of Technology, Graduate School, ATLANTA, GA. 1 Niche users give it an average review of 5 stars. Featured Review: Other says Georgia Tech is among the top colleges you can attend. My journey there had its highs and lows, but looking back, every experience contributed significantly to my growth. A piece of advice I'd offer... Read 1 reviews.

College of Arts and Sciences - Lehigh University

• Lehigh University ·
• BETHLEHEM, PA

College of Engineering and Science at Florida Institute of Technology

• Florida Institute of Technology ·

California State University - East Bay

• HAYWARD, CA
• · Rating 4.42 out of 5   50

Showing results 1 through 25 of 398

Ph.D. Program in Biology

Our graduate program provides students of diverse backgrounds with the opportunity to intensely engage in research in the biological sciences and contribute to the broader scientific community.

Average time to degree: 5.3 years

Percentage of graduates in post-doctoral or permanent positions: 94%

Average publications after completion of program: 3.8 per person

Ph.D. Program

The Department of Biology introduces graduate students to diverse fields of biological science, and provides them with expert guidance to excel in research. The department is invested in training students to become excellent scientists, researchers, science communicators, and instructors. We are a diverse and global community, committed to expanding scientific career opportunities to all. Some of our graduates become academics, whereas others find careers in government, private industry, public policy, or elsewhere (see where recent graduates are now). The ability to communicate ideas and research results clearly and convincingly is key to success in any career.

Experience Biology at Georgetown University

Professors and current students share their perspectives on Georgetown’s cutting-edge biology graduate program.

Potential applicants are urged to identify and contact potential research mentors directly before applying. Please refer to the research page and the list of faculty interested in accepting students . Not all laboratories will have open positions available for a new graduate student in a given year.

Financial Assistance for PhD Students

    students who are accepted into the phd program in biology are guaranteed a minimum of five years of full funding. this funding includes:.

Stipend The stipend for PhD students in Biology is set by the Collective Bargaining Agreement between Georgetown University and the Georgetown Alliance of Graduate Employees ( GAGE ). The current stipend rate is $36,934 for the year. The stipend is paid on a 12-month contract, typically in 26 biweekly paychecks.

Tuition Waiver The Graduate School of Arts & Sciences (GSAS) provides full tuition waivers for all PhD students for the duration of the PhD program.

Health Insurance Health insurance is required. The GSAS provides free student health insurance to all students. The insurance covers doctor’s visits, hospital stays, and prescription drugs. Students may waive this benefit if they prefer a different plan through parents or a spouse.

Dental Insurance The GSAS provides free dental insurance to all students. The insurance covers routine dental care, such as cleanings and exams.

Yates Athletic Facility All students are eligible to use the Yates athletic facility for free. The facility has a gym, swimming pool, and fitness classes.

Parental Leave Graduate student workers are guaranteed six weeks of paid parental leave. This leave can be used to bond with a new child or to care for a sick child.

Medical Leave Graduate student workers are guaranteed six weeks of paid medical leave. This leave can be used to recover from a medical condition or to care for a sick family member.

To learn more about financial assistance for PhD students, please visit the Graduate Funding page. You can read more about what it means to do a PhD in a union-protected graduate program  here

Student Quotes

“Until you start your journey in grad school you might not know what you are missing out on. Other than the course work, grad school has been instrumental in  inculcating important life values and skills such as confidence, time management, survival, and holding personal and professional relationships in higher esteem. ”

“Studying at Georgetown has enabled me to be a lot more confident in my ability to ask questions and chart my own path in scientific research. My advisor and mentors here at Georgetown have been top quality in terms of both academic and personal support”

“Georgetown is a place with an environment conducive to learning and professional growth for prospective scientists working towards their graduate degrees and beyond. In this program, I found mentorship and training that foster the critical thinking required to answer the questions I am interested in. I found the multidisciplinary nature of the graduate student body to be engaging and supportive from the beginning of my journey.”

“The breadth of research that goes on in the Bio department really facilitates a broad understanding of biology from the life of single cells to the life of much larger organisms. It really is something unique that you don’t get at other more specific departments and has really helped me think outside of the box with my own work.”

Graduate Research Opportunities

Learn about the research programs that the department of Biology offers and find out which programs are recruiting graduate students.

Graduate Teaching Opportunities

Find out about the many opportunities that are offered to graduate students to develop teaching and mentoring skills.

Graduate Funding Opportunities

Learn about stipend funding and the additional funds from the GU Graduate School offered to help support graduate dissertation research or travel to meetings.

Biological and Biomedical Sciences

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Biological and Biomedical Sciences (BBS)  is an area of study within the Division of Medical Sciences, an administrative unit based at Harvard Medical School that coordinates biomedical PhD activities at the Longwood Medical Area. Students who study in BBS receive a PhD in medical sciences. Prospective students apply through the Harvard Kenneth C. Griffin Graduate School of Arts and Sciences (Harvard Griffin GSAS). In the online application, select  “Division of Medical Sciences” as your program choice and select "Biological and Biomedical Sciences" in the area of study menu.

This program trains you in the biosciences starting with core training in contemporary genetics, biochemistry, and molecular, cellular, and mechanistic biology. You can customize your curriculum to align with your research interests.

You will have access to a vast wealth of resources, including more than 700 faculty, top scientists, core facilities and hospitals, cutting-edge research, and an extensive alumni network. You will be in close proximity to top nationwide hospitals and research centers and more than 130 biotech, biopharma, and pharmaceutical companies that have biological and biomedical sciences (BBS) alumni on staff.

Examples of specific student projects include “Development of nucleic acid detection methods for object provenance and viral diagnostics,” “Interrogating genetic diversity in Mycobacterium abscessus with transposon-sequencing,” and “Sexual dimorphism in hair follicle stem cells.”

Graduates of the program have secured faculty positions at such prestigious institutions as New York University, University of Minnesota, Harvard University, and University of Illinois. Others have gone on to careers with leading companies such as Novartis and the Broad Institute. Derrick Rossi, cofounder of Moderna, is a graduate of the BBS program.

Personal Statement

Standardized tests.

GRE General: Not Accepted GRE Subject: Not Accepted iBT TOEFL minimum score: 100 IELTS minimum score: 7

See list of Biological and Biomedical Sciences faculty

APPLICATION DEADLINE

Questions about the program.

Graduate Admissions

Helpful guides for applying to the biology phd program, tips for applying to the cmob track, tips for applying to the eco/evo track.

The PhD program trains students for careers in research and teaching. In addition to a wide range of courses, the PhD program offers informal but intensive instruction in laboratories and seminars, characterized by close contact between students and their faculty advisors. Biology is one of fourteen home programs  in the  Biosciences,  which span the School of Medicine and the School of Humanities and Sciences.

How to Apply

The 2024-2025 Biology PhD application processes is run through strictly through Biology (School of Humanities & Sciences). 

Previously part of the Biology Department , Hopkins Marine Station is now part of the Oceans Department within the Stanford Doerr School of Sustainability (SDSS).  Several faculty took appointments in Oceans, while others chose to stay in Biology; still others have joint appointments.  Additionally, several faculty members in Ecology/Evolutionary Biology now have joint appointments within Biology and SDSS. Based on these changes, the following information will provide further guidance for the application process. 

• Faculty with 100% appointments in Biology, will only recruit through Biology.  The Biology application will be used.
• Ecology/Evolutionary Biology faculty with 50/50 appointments in both Biology and SDSS will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Ecology/Evolutionary faculty with joint Biology/SDSS appointments, you should reach out to them directly regarding the appropriate application track to choose. 
• 100% Oceans faculty will only be recruiting students through the Oceans application.
• Oceans faculty appointed 50/50 in Biology and SDSS (Oceans Department), will recruit students through either Biology or Oceans .  If you are interested in working with Oceans faculty with joint Biology/SDSS appointments, you should reach out to the individual faculty members you hope to work with regarding the appropriate application and track to choose. 

Faculty Affiliated with the Stanford Doerr School of Sustainability

Oceans Department:

Fio Micheli 100% 

Jeremy Goldbogen 100%

Giulio De Leo 100%

Larry Crowder 100%

Steve Palumbi 50% Oceans/50%Biology

Barbara Block 50% Oceans/50% Biology

*Note Chris Lowe is 100% in Biology

Ecology/Evolution

Liz Hadly 50% SDSS/50%Biology

Rodolfo Dirzo 50% SDSS/50%Biology

Tad Fukami 50% SDSS/50%Biology

Kabir Peay 50% SDSS/50%Biology

Chris Field 50% SDSS/50% Biology

Gretchen Daily 50% SDSS/50% Biology

Application and Deadline

Stanford Biology PhD Program applications are made through Graduate Admissions . The application deadline for Autumn Quarter 2025 matriculation is December 3, 2024 at 11:59pm pst.  The application for the Autumn 2025 cohort will be available in September 2024.  Please review the Graduate Admissions website prior to starting your application. Applications are considered for admission in Autumn Quarter only. 

Application Status

To check your application status,  click here to Visit Your Status Page .  

Requirements

• U.S. Bachelor's degree or its foreign equivalent
• Stanford Graduate application
• Non-refundable application fee of $125 (or fee waiver found here ) 
• Statement of Purpose
• Three Letters of Recommendation (a maximum of four letters will be accepted)
• Transcripts - Unofficial transcripts from all universities and colleges you've attended for one year or more must be uploaded to the online application.  Graduate Admissions only requires admitted applicants who accept the offer of admission to submit official transcripts that shows their degree conferral. Please do not send or have sent any official transcripts to us at this time.
• TOEFL Test of English as a Foreign Language (required for non-native English speakers)
• A current Curriculum Vitae/Resume

Knight-Hennessy Scholars

Join dozens of  Stanford School of Humanities and Sciences students  who gain valuable leadership skills in a multidisciplinary, multicultural community as  Knight-Hennessy Scholars  (KHS). KHS admits up to 100 select applicants each year from across Stanford’s seven graduate schools, and delivers engaging experiences that prepare them to be visionary, courageous, and collaborative leaders ready to address complex global challenges. As a scholar, you join a distinguished cohort, participate in up to three years of leadership programming, and receive full funding for up to three years of your PhD studies at Stanford. Candidates of any country may apply. KHS applicants must have earned their first undergraduate degree within the last seven years, and must apply to both a Stanford graduate program and to KHS. Stanford PhD students may also apply to KHS during their first year of PhD enrollment. If you aspire to be a leader in your field, we invite you to apply. Learn more about  KHS admission .

Biology PhD Program General Information

Diversity, inclusion, equity, and belonging.

The Department of Biology recognizes that the Supreme Court issued a ruling in June 2023 about the consideration of certain types of demographic information as part of an admission review. All applications submitted during upcoming application cycles will be reviewed in conformance with that decision. The Department of Biology welcomes graduate applications from individuals with a broad range of life experiences, perspectives, and backgrounds who would contribute to our community of scholars. Review of applications is holistic and individualized, considering each applicant’s academic record and accomplishments, letters of recommendation, and admissions essays in order to understand how an applicant’s life experiences have shaped their past and potential contributions to their field.Please see the websites below for more resources.

• Diversity and Engagement in Biosciences PhD Programs
• Vice Provost for Graduate Education
• Biology Equity and Belonging
• Research Sites
• Faculty Profiles and Labs
• Inquiries about the PhD in Biology should be directed to  biologyadmissions [at] lists.stanford.edu (biologyadmissions[at]lists[dot]stanford[dot]edu)

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Biology, PhD

The Biology Graduate Program represents many areas of biology, and interactions with a diverse group of colleagues provide opportunities to broaden every student’s thinking and make connections between different fields and scientific approaches. Areas of research include microbiology, cell biology, development, physiology, neuroscience, animal behavior, plant biology, genetics, computational biology, evolution, ecology and biodiversity. 

Each entering graduate student has the freedom to pursue topics ranging from the behavior of molecules to that of cells, organisms, genomes, and ecosystems. We encourage students to get broad exposure through lab rotations with any faculty member in the Biology Graduate Group. As students focus on more specific research interests, they tailor their graduate education accordingly, choosing courses from different departments and schools at Penn as appropriate.

Students complete most of their course work and lab rotations in the first year and then start their thesis research in the second year while completing their teaching requirement and preparing for their candidacy exams.  Students are then fully focused on thesis research by the end of the second year.  Students still have the option of taking additional courses in advanced years in order to enhance their graduate research.

For more information: http://www.bio.upenn.edu/graduate/

View the University’s Academic Rules for PhD Programs .

Required Courses

The total course units for graduation in this program is 13.5.

See the website for a list of electives:  http://www.bio.upenn.edu/graduate/handbook/academic-topics/course-requirements

The degree and major requirements displayed are intended as a guide for students entering in the Fall of 2024 and later. Students should consult with their academic program regarding final certifications and requirements for graduation.

Sample Plan of Study

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Graduate program.

Duke Biology offers diverse training opportunities for students seeking a Ph.D. degree in biological sciences. Our program is highly flexible, allowing students to design the path that best fits their professional needs. Our students spend the majority of their time engaged in innovative research, and have the opportunity to interact and collaborate with researchers from a wide range of fields. 

The graduate students in the Biology Department (Biograds) are integral to the department and community. Contributions include: running weekly happy hours and prospective student recruitment weekends as well as organizing seminars, journal clubs, teaching-assistant training orientation, and diversity-themed workshops.

Program Requirements

Requirements for the Ph.D. include course work, participation in seminars relevant to your field of research, successful completion of an oral qualifying examination, one year of teaching experience, completion and defense of a research dissertation under faculty guidance.

Affiliated Graduate Programs

Duke Biology is involved in multiple University-wide graduate programs, which link researchers across different departments and schools at Duke – most notably the basic sciences departments in the  Duke University Medical Center  and the  Nicholas School of Earth and Ocean Sciences .

For Prospective Students

Learn more about our degree, our university, and our community. We'll explain the application process, financial support, and more.

• For Current Students

We've compiled resources to assist you while working toward your Ph.D. – from requirements and fellowships, to milestones, useful Duke links, and more.

Learn more about the application process, deadlines and other resources for our candidates.

Review Application Information

• Duke Biology’s Mission Statement
• AJED Annual and Semester Reports
• AJED Meeting Notes
• Biology Cultural Association (BCA)
• Inclusion, Diversity, Equity, and Antiracism Committee (IDEA)
• Learning from Baboons: Dr. Susan Alberts
• Extremophiles and Systems Biology: Dr. Amy Schmid
• How Cells Manage Stress: Dr. Gustavo Silva
• Predator-Prey Interactions in a Changing World: Dr. Jean Philippe Gibert
• Exploring the Extracellular Matrix: Dr. David Sherwood
• Cell Division's Missing Link: Dr. Masayuki Onishi
• Listening in to Birdsong: Dr. Steve Nowicki
• Biogeochemistry as Ecosystem Accounting: Dr. Emily Bernhardt
• Building a Dynamic Nervous System: Dr. Pelin Volkan
• Investigating a Key Plant Hormone: Dr. Lucia Strader
• Imagining Visual Ecology: Dr. Sönke Johnsen
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